Ok, actualy your long post helps alot figuring out whats wrong with him. And good job nursing the poor guy back to health, have done it a few times and its a very rewarding experience. Back to topic, considering what your telling me I ask your igs diet, temps and expossure to direct sun ligth or uv lamp (also wich brand uv bulb do you use and how long has it been since you last replaced it), so we can come up with a good gess

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