According to this guy:
www.tortoisetrust.org/articles/webdiet.htm

His mantra in this article is if you would never find a food item in the natural environment of a tortoise, then you need to exclude or limit that food item in the captive diet. I think that is great advice about all herbivores. Now there are some things we can substitute for things we can't get for our captive lizards, but staples should be low in sugar, low in protein, low in oxalates,and lower in water content than most of the greens sold at super markets, and they should be high in fiber.

On the sugar, I think it causes diarrhea in herbivores, but I don't have time right now to look that up. You might find the answer in the link I posted above.

I am not going to respond to questions about diet again in this thread because it is really an enclosure thread. We can start another to discuss diet.

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