Field study methods and husbandry are normally based on a snake, an individual snake. That is, follow a snake. Put a snake in a cage. All about one snake. In the past science has treated accepted these animals as solitary, as you do with your moss cage. Most do, so don't feel bad.
The paper you cited, points out that the biological development, in particular, the system of pheromones, indicates a society of snakes. or better yet, a community of snakes. It indicates advanced biological systems to communicate with other reptiles. Not only their own species by the way.
What is interesting is me is, the balance of pheromones, vs. cognitive choice(which they are not suppose to have but do have) That is, when a individual snake behaves in a way, contrary to pheromone exposure. And they do. After all, is a language. You tell me to go away and I don't go. Why, because I don't want to. So what happens when this occurs in nature or our cages. Dude, this is so much fun its ridiculous.
If you thought you could not shut me up before, get ready, cause on this subject, I could go on forever. Thank you so much for bringing this subject up. Are you sure you want me to be part of your thread, because I am only getting started here, I can and will go on and on and on. Love it. I do.

A result of pheromone conflict with civilized animals. A non violent battle. I love it, they are more civilized then us. hahahahahaha

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