I have bred normal looking Harlequins to unrelated animals,and produced offspring that could certainly pass for first-rate Jungles...can the Harlequin influence be "bred out"? in all honesty it has never been a goal,so I can't say I've persued that thought to its extreme end. I think you're kind of mistaken in your thought that Harlequins were simply line bred for aberrancies...many of the key breedings I focused on involved non-abberant animals from this line,that were selected because of their parentage rather that their appearence.
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