Bryant did prove his Ghost to be hypomelanistic when he bred it to a normal Nicaraguan, but the Type II part needed to be proof. Do not quote me on this, but I am almost certain Gus' Hypo that he bred came from offsping of that Ghost. While there were a few Nicaraguan Ghosts floating around before Gus' litter, they were farmed imports.
Gus was the first person to produce neonatal Ghosts in the United States through his own captive breeding program (and it was an accident).

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