Ball Python genetics, as well as some boa genetics are
"ass-backwards" of recessive colubrid genetics.
Melanistic(hypermelanism)genes are co-dominant in corns along with the reds......melanophores produce the dark pigment,....and hypermelanistic animals possess an extensive amount of these melanophores.
The red/orange coloration is produced from Xanthophores. These chromatophores produce reds,orange and yellows, and in cornsnakes, can produce intermediate shades of these colors as well, but have a predominantly red coloration from cells known as "erythrophores". Cornsnakes usually have a tendency to become more yellow as they age, especially in the first third of their body and is thought to be due to "carotenoid" retention from their diet.
Since the dark pigment on the "hypermelanistic" animal is a wild-caught "normal" gene, just a greater amount of the cells that produce dark coloration as mentioned above, it wouldn't necessarily be expressed with any certainty to the offspring like it absolutely would in a amelanistic pairing, or many other recessive pairings for that matter.
I'm not trying to come off as some sort of genetisist to show-off all the big words,.....just trying to help explain what is responsible for what!LOL
So in the hypothetical(another big word!LOL) situation you mentioned before, the snakes that would be bred would'nt be anything more than leucistic, or het for leucistic animals with a normal wild-caught predominance for being darker animals.
If you had TWO very dark corns in a pairing, you would have a tendency to produce darker than normal offspring.
This will either help, or possibly confuse you even more!LOL, but I hope it helps.
best regards, ~Doug
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"Better to be silent and thought a fool, than to open mouth and remove any doubt!"