When learning genetics, you need to understand that genes come in pairs. When the pair of genes can be selected from two different genes (call them A and a), there are three possible combinations:
AA = homozygous for A
Aa = heterozygous
aa = homozygous for a
When making sex cells (sperm or eggs), each sex cell gets only one of each pair of genes. So a snake that is AA only produces sex cells that are A, a snake that is aa only produces sex cells that are a, and a snake that is Aa produces sex cells with half of them being A and half of them being a.
When the sperm fuses with the egg, the result has a pair of genes again.
It might help to take some scraps of paper and write A on some and a on others. Then play around making combinations of two scraps.
When there are two possible genes to select from, there are six possible matings:
1. AA x AA - both make only A sex cells.
Result: all babies are AA.
2. AA x Aa - The AA parent makes only A sex cells. The Aa parent makes 1/2 A sex cells and 1/2 a sex cells.
Result: A joins with 1/2 A to make 1/2 of the babies AA.
A joins with 1/2 a to make 1/2 of the babies Aa.
3. AA x aa - The AA parent makes only A sex cells. The aa parent makes only a sex cells.
Result: all babies are Aa
4. Aa x Aa - both parents produce 1/2 A sex cells and 1/2 a sex cells. This is the tricky one.
Result: 1/2 A - 1/2 A makes 1/4 AA
1/2 A - 1/2 a makes 1/4 Aa
1/2 a - 1/2 A makes 1/4 aA
1/2 a - 1/2 a makes 1/4 aa
As gene order on paper makes no difference, Aa and aA are the same thing. The final result is
1/4 AA
2/4 Aa
1/4 aa
5. Aa x aa - The aa parent makes only a sex cells. The Aa parent makes 1/2 A sex cells and 1/2 a sex cells.
Result: a joins with 1/2 A to make 1/2 of the babies Aa.
a joins with 1/2 a to make 1/2 of the babies aa.
6. aa x aa - both make only a sex cells.
Result: all babies are aa.
Let's assign a to the piebald gene and A to the normal gene. A snake that is piebald is aa, a normal snake is AA, and an Aa snake is heterozygous piebald and looks normal. A normal to piebald mating is mating 3 above, and a het piebald to het piebald mating is mating 4 above.
And 100% het piebald simply means certainty that a normal looking snake is actually het piebald. Such snakes are produced in matings 3 and 5. Het piebalds are also produced in matings 3 and 4, but they are mixed with normals. In matings 3 and 4, there is a less than 100% chance that a given normal-looking baby snake is actually het pied.
Paul Hollander