your pretty spot on malnutrition and bad sheds could have caused it. the sheds usually calim toenails and occasionally toes (but bad sheds are atributed to low humidity) the handling could also have caused this. its not certain but another consideration is that maybe it was kept with more than one at one time and was bitten by a more agressive cage mate lol i dont know much of the situation but im glad its in good hands now
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REAPER REPTILES