Perry,
The pushups can definitely be associated with breeding behavior, but the two don't always go hand in hand. The pushup is definitely more of a broad statement, than just stating the desire to mate. I would think of it as an advertisement that large, dominant males make, to show that they are in charge of a certain territory. This may warn other males to stay away, as well as tell females that they may have to put up with his breeding attempts as long as she is in his territory. Females will also do the pushups, but there seems to be a subtle difference in technique. The female's pushup could be more of a submissive message, or even one of defiance, if she is unready to breed. We have found some of our pairs in the act, but have not witnessed much of the preceeding behavior.

Nick

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