Hi Arron, Its not an A or B type of answer. First, eggs are very tolerant. After all, they squeeze out the females bum. Have you see that?
And yes, the zygote is located in the middle of the egg before being laid and after its laid, the zygote migrates to the upper part of the egg.
It appears there is a critical time when the eggs can fail if rotated. I do not know how long that is. When first laid, the eggs can be rolled tossed around, etc. Then later in development, they can be rolled etc etc, many stories, like my pet ferret got to some cornsnake eggs and moved them to its cage and they still hatched. Even though many had tooth holes in them.
I think the critical time is withing the first couple of weeks.
Lastly, I have moved them even at that time with no problems. I guess we have to define what moving them means and who had how that rule was made. For instance, rotating weak/dead eggs is a problem, rotating healthy eggs may not be. Simple put, I do not worry about it, but I don't rotate them for fun.

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