Another question I have is, why do you say there is no gene flow between populations in getula? I can understand no gene flow between montanes and other species that are isolated but with lower altitude generalists like kings and gophers, etc. not so much. Even though they form family groups they still overproduce and that would result in some leaving the "family" of their birth/hatching and possibly assimilating into another "family". There is nothing biological that would totally prevent Cals from mixing with splendida/mbk, the only thing that could prevent it would be environmental factors and those don't seem to present a total barrier since they can inhabit so many types of environments.
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