Hi, When your talking behavior(ethology) you need to understand what that means. First it means, tendencies and trends. It does not ever mean All and always.

With animals,(and people) social training occurs at birth. Any social animal raised alone, becomes socially problematic. Again, including people. So your first mistake is, you raised them alone. Therefore, you have no reason to ever think they will socialize normally. Back to behavior, they still can. Isn't behavior cool? Behavior has soft rules, not hard fast rules.

When raised with other animals, its more then simple bonding with a mate. IT teaches them how to live with other individuals and WHICH other individuals to trust. Out of that, normally they pick a mate to bond with.
When they Bond with a mate they were raised with, they are totally trustworthy. Two individuals that you introduced at a late age, they do not bond to the same degree, they may breed and not eat eachother, yet. Pairs introduced in the beginning of winter, are more trustworthy then those introduced in spring, etc etc. Folks its all by degree.

So can they be kept together without bonding, yes sometimes. But that's not the problem. The problem is, when they do not tolerate eachother, like your experiencing now. If they are normal, the above applies, end part 1

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